1Laboratoire d'Océanographie de Villefranche, Sorbonne Université, CNRS, F-06230 Villefranche-sur-Mer, France; email: [email protected][email protected][email protected]2Advanced Science-Technology Research (ASTER) Program, Institute for Extra-Cutting-Edge Science and Technology Avant-Garde Research (X-STAR), Japan Agency for Marine-Earth Science and Technology, Yokosuka, Kanagawa 237-0021, Japan; email: [email protected]3School of Marine Sciences, University of Maine, Orono, Maine 04469, USA; email: [email protected]